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Development of a Plant Shoot Temperature Model for Greenhouse Climate Management Progress Report — September 1996

Date Aug. 26, 1996

Title of Project Control of tomato spotted wilt virus using

transgenic plants that produce virus-specific monoclonal antibodies

Institution where work is being conducted University of Hawaii

Amount of Endowment Grant $25,000

Covering Period Jan. 96 to Dec. 96

Anticipated Date of Project Completion/Final Report 1997

Individual(s) Conducting Project:

(List Project Leader First)

John S. Hu - Title Assistant Professor

Telephone Number (808) 956-7281

M. Q. Xu - Title Graduate Assistant

Control of Tomato Spotted Wilt Virus Using Transgenic Plants that Produce

Virus-Specific Monoclonal Antibodies

John S. Hu and M. Q. Xu

University of Hawaii

Progress Report to the American Floral Endowment, 8/26/96
A. Project Objectives:

Engineer and express in plants the genes of the monoclonal antibody which

reacts to tomato spotted wilt virus (TSWV).

B. Summary of work:

Hybridoma cell lines secreting monoclonal antibodies to TSWV have been

produced. One cell line was selected for the cloning of the antibody genes.

Universal degenerate primers were designed for PCR amplification of the

variable DNA fragments of heavy and light chains of monoclonal antibodies.

The PCR products were ligated into one single-chain Fv antibody gene construct

and transformed into tobacco Nicotiana tabacum. Some of the transgenic

tobacco plants were resistant to systemic infection of TSWV. The transgenic

lines were also characterized by Southern and Northern blot hybridizations.

C. Results to date:

Hybridoma cell lines producing specific monoclonal antibodies to TSWV have

been made. One cell line (TSWV-MAb8C4D6), which has broad specificity to

TSWV isolates and reacts to the nucleoprotein of TSWV, has been selected

for the cloning of the antibody genes. Universal degenerate primers were

designed for amplification of variable regions of heavy and light chains

of monoclonal antibodies. PCR products were ligated into one single-chain

antibody gene construct and then cloned into a plant transformation vector,

and used to transform tobacco. Transgenic plants have been produced and

characterized. Several lines showed clear delay in symptom development.

Two of the transgenic tobacco lines were completely resistant to systemic

infection of TSWV. These lines were tested more than five times and

identical results were obtained. Progenies from the two resistant lines

were also resistant to systemic infection of TSWV. Southern and Northern

blot hybridization analyses were done with the transgenic plants. The RNA

expression level of the transgenic plants correlate with the virus resistance

level. Specific antibodies were produced against the single-chain antibody

and will be used in Western blot analysis to examine the expression of

the antibody protein in the transgenic plants.

D. Future plans covered by the Endowment grant:

The resistant transgenic plants will be examined in Western blot hybridization

and for resistance to other tospoviruses.

E. Anticipated benefits for floral industry:

It was recently reported that transgenic plants expressing a monoclonal

antibody against the coat protein of a tombusvirus have been produced.

Their data show a delay in symptom development and reduction on virus replication

suggesting a role of the antibodies in plant protection. The long term

goal of this research is to control TSWV using transgenic plants that produce

TSWV-specific monoclonal antibodies. Since TSWV has a very wide host range,

infecting 192 dicotyledonous species in 33 families and eight monocotyledonous

species in 5 families. If this approach works, the specific genes that

encode monoclonal antibodies to TSWV could be introduced into many floricultural

crops, for control of this devastating virus disease. Currently, several

other labs are evaluating this antibody-mediated protection strategy for

control of different plant viruses.