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Control of Tomato Spotted Wilt Virus Using Transgenic Plants that Produce Virus-Specific Monoclonal Antibodies Progress Report — September 1993

Date 8/23/93

Title of Project Control of Tomato Spotted Wilt Virus Using Transgenic Plants that

Produce Virus-Specific Monoclonal Antibodies

Institution where work is being conducted University of Hawaii

Amount of Endowment Grant $ 10,000
Covering Period Jan. ‘93 to Dec. ‘93

Anticipated Date of Project Completion/Final Report 1996

Individual(s) Conducting Project:

(List Project Leader First)

Dr. John S. Hu - Title Assistant Professor

Telephone Number (808) 956-7281

Mr. Z. C. Wu - Title Graduate Assistant

Control of Tomato Spotted Wilt Virus Using Transgenic Plants that Produce Virus-Specific Monoclonal Antibodies

John S. Hu and Z. C. Wu

University of Hawaii

Progress Report to the American Floral Endowment, 8/23/93

A. Project Objectives:

1) Clone the genes of the Ig gamma and kappa protein chains of the monoclonal

antibody which reacts to TSWV.

2) Sequence and engineer the genes and subclone the genes into expression

vectors.

B. Summary of Work Conducted:

MAb’s specific to TSWV have been characterized in ELISA and Western blot

analysis. One hybridoma cell line (TSWV-MAb 8C4D6) has been selected for the

cloning of the antibody genes. Full length RNAs that code for the Ig gamma and

kappa proteins were used for cloning. Complementary DNAs were produced to these

RNAs using oligo-dT as a primer and reverse transcriptase. Specific clones identified

in colony hybridization have been characterized.

C. Results to Date:

Hybridoma cell lines producing specific monoclonal antibodies to TSWV have

been made. One cell line (TSWV-MAb8C4D6), which has broad specificity to

TSWV isolates and reacts to the nucleoprotein of TSWV, has been selected for the

cloning of the antibody genes. It has been determined that the monoclonal antibody

has a gamma 1 heavy chain and kappa light chain. RNAs have been isolated from

the cell line for cloning of the antibody heavy chain and light chain genes. Specific

oligonucleotide probes to the conserved regions of the genes have been made. Full

length mRNAs that code for the Ig gamma and kappa proteins were used for cloning.

Complementary DNAs were produced to these mRNAs using oligo-dT as a primer

and reverse transcriptase. Specific clones identified in colony hybridization have been

sequenced. The clones were verified by checking the sequences against

published sequences of the conserved and the leader sequence regions.

Specific primers were designed to amplify some frangments of the

genes in polymerase chain reaction to obtain full length clones.

D. Future Plans Covered by the Endowment Grant:

Clones containing the complete genes of gamma I heavy chain or kappa light

chain and the leader sequences will be engineered for transformation work.

E. Anticipated Benefits for Floral Industry:

Tomato spotted wilt virus (TSWV) is one of the most devastating diseases on

floricultural and vegetable crops in Hawaii and worldwide. This virus disease is very

difficult to control. Recent developments in biotechnology have provided new

opportunities to solve practical agricultural problems. Genetic engineering offers new

approaches to produce virus resistant varieties. A recent scientific breakthrough has

presented a new possibility for controlling plant virus diseases through the use of

transgenic plants that produce antibodies to specific plant viruses. It was recently

reported that transgenic plants expressing a monoclonal antibody

against the coat protein of a Tombusvirus have been produced

(Tavladoraki et al. 1993). Their preliminary data show a delay in

symptom development suggesting a possible role of the antibodies in

plant protection. The antibody molecules may bind to the nucleoproteins to

prevent uncoating in the early stage of infection, or bind to the nucleoprotein

molecules to prevent assembly of virions in the later stages of virus replication. Such

a system would be analogous, in a general way, to the common antibody defense

system in animals. The long term goal of this research is to control TSWV using

transgenic plants that produce TSWV-specific monoclonal antibodies. Since TSWV

has a very wide host range, infecting 192 dicotyledonous species in 33 families and

eight monocotyledonous species in 5 families. If this approach works, the specific

genes that encode monoclonal antibodies to TSWV could be introduced into many

floricultural crops, for control of this devastating virus disease.